A list of publications from the BEHAVE project can be found at the USU Library in the Digital Common.

Citations are listed by year and they include abstracts and in some cases the complete article.

Behave Facts

Another way to look at our publications and their major findings is to view the Behavior Facts page. It lists major findings of the BEHAVE Project and references. It is current through 2006, but will soon be updated.

Recent Publications


1.   Mote, T.E., J.J. Villalba, and F.D. Provenza. 2007. Relative availability of tannin- and terpene-containing foods affects food intake and preference by lambs. J. Chem. Ecol. 33:1197–1206.

2.   Papachristou, T.G., L.E. Dziba, J.J. Villalba, and F.D. Provenza. 2007. Patterns of diet mixing by sheep offered foods varying in nutrients and plant secondary compounds. Appl. Anim. Behav. Sci. 108:68-80.


1.   Dziba, L.E. and F.D. Provenza. 2008. Dietary monoterpene concentrations influence feeding patterns of lambs. Appl. Anim. Behav. Sci. 109:49–57.

2.   Lyman, T.D. F. D. Provenza and J. J. Villalba 2008. Sheep foraging behavior in response to interactions among alkaloids, tannins and saponins. Journal of the Science of Food and Agriculture 88:824–831.

3.   Mote, T., J.J. Villalba, and F.D. Provenza. 2008. Foraging sequence influences the ability of lambs to consume foods containing tannins and terpenes. Appl. Anim. Behav. Sci. 113:57–68.

4.   Rogosic, J., J.A. Pfister, F.D. Provenza, and J. Pavlicevic. 2008. The Effect of Polyethylene Glycol on Intake of Mediterranean Shrubs by Sheep and Goats. Journal of Animal Science 86:3491-3496.

5.   Villalba, J.J., F.D. Provenza, and J.O. Hall. 2008. Learned appetites for calcium, phosphorus, and sodium in sheep. J Anim Sci. 86:738-747.


1.   Lisonbee, L.D., J.J. Villalba, F.D. Provenza, and J.O. Hall. 2009. Tannins and self-medication: Implications for sustainable parasite control in herbivores. Behav. Process. 82:184-189.

2.   Lisonbee, L.D., J.J. Villalba, and F.D. Provenza. 2009. Effects of tannin on selection by sheep of forages containing alkaloids, tannins and saponins. J. Sci. Food Agric. 89:2668-2677.

3.   Mantz, G. J. J. Villalba and F. D. Provenza. 2009. Supplemental polyethylene glycol affects intake of and preference for sericea lespedeza. J. Anim. Sci. 87:761-769.

4.   Villalba, J.J., X. Manteca and F.D. Provenza. 2009. Relationship between reluctance to eat novel foods and open-field behavior in sheep. Physiol. Behav. 96:276–281.

5.   Villalba, J. J., Provenza, F. D., & Stott, R. 2009. Rumen distension and contraction influence feed preference by sheep. Journal of Animal Science, 87:340-350.


1.   Catanese, F. R. A. Distel, R. M. Rodrguez-Iglesias and J. J. Villalba. 2010. Role of early experience in the development of preference for low-quality food in sheep. Animal 4:784–791.

2.   Villalba, J.J, F.D. Provenza, J.O. Hall and L.D. Lisonbee. 2010. Selection of tannins by sheep in response to gastrointestinal nematode infection. J. Anim. Sci. 88:2189-2198.


1.   Sanga, U., F.D. Provenza, and J.J. Villalba. 2011. Transmission of self-medicative behaviour from mother to offspring in sheep. Animal Behaviour 82:219-227.

2.   Lyman, T.D., F.D. Provenza, J.J. Villalba and R.D. Wiedmeier. 2011. Cattle preferences differ when endophyte-infected tall fescue, birdsfoot trefoil, and alfalfa are grazed in difference sequences. Journal of Animal Science 89:1131–1137.

3.   Bach, A., J.J. Villalba, and I. R. Ipharraguerre. 2011. Interactions between mild nutrient imbalance and taste preferences in young ruminants. Journal of Animal Science 90:1015-1025.

4.   Villalba, J.J., A. Bach, and I. R. Ipharraguerre. 2011. Feeding behavior and performance of lambs are influenced by flavor diversity. Journal of Animal Science 89:2571-2581.

5.   Villalba, J.J., F.D. Provenza, A.K. Clemensen, R. Larsen, and J. Juhnke. 2011. Preference for diverse pastures by sheep in response to intraruminal administrations of tannins, saponins, and alkaloids. Grass Forage Sci. 66: 224–236.


1.   Catanese, F., R. A. Distel, F. D. Provenza, and J. J. Villalba. 2012. Early experience with diverse foods increases intake of nonfamiliar flavors and feeds in sheep. J. Anim. Sci. 90:2763-2773

2.   Juhnke, J., J. Miller, J.O. Hall, F.D. Provenza, and J.J. Villalba. 2012. Preference for condensed tannins by sheep in response to challenge infection with Haemonchus contortus. Veterinary Parasitology 188:104-114.

3.   Lyman, T.D., F.D. Provenza, J.J. Villalba, and R.D. Wiedmeier. 2012. Phytochemical complementarities among endophyte-infected tall fescue, reed canarygrass, birdsfoot trefoil and alfalfa affect cattle foraging. Animal 6: 676-682.

4.   Owens, J., F.D. Provenza, R.D. Wiedmeier, and J.J Villalba. 2012. Influence of saponins and tannins on intake and nutrient digestion of alkaloid-containing foods. Journal of the Science of Food and Agriculture, 92: 2373-2378.

5.   Owens, J., F.D. Provenza, R.D. Wiedmeier, and J.J. Villalba. 2012. Supplementing endophyte-infected tall fescue or reed canarygrass with alfalfa or birdsfoot trefoil increases forage intake and digestibility by sheep. Journal of the Science of Food and Agriculture 92:987-992.


Jensen, T.L., F.D. Provenza and J.J. Villalba. 2013. Influence of diet sequence on intake of foods containing ergotamine d-tartrate, tannins and saponins by sheep. Applied Animal Behaviour Science 144:57-62.